During the storage of grains, various biotic factors such as insects, rodents, mites, fungi, bacteria, etc. and various abiotic factors such as inappropriate moisture, temperature, lack of sunlight, flood, etc. are responsible for losses of grains. These factors act on stored grains and result in degradation, poor germinability, discoloration, etc. Both the factors lead to the spoilage and wastage of seeds and grains. Hence all of them are correct.